limnology syllabus

 
4 cr. Spring 2001

 

Instructor:      John E. Havel, Ph.D.
                        Office:  KGSX 307      phone: 836-5308  email: johnhavel@missouristate.edu
                        Office hours: MW 10-11, F 10-12, & by appointment

 

Course content: Limnology is the study of inland waters, and has as its central focus the structure and function of lakes.  Limnology is multidisciplinary, with topics including lake formation, light and heat, hydrology, water chemistry, plant and animal diversity, and the ecology of food webs. BIO 562 emphasizes the laboratory and will include one weekend field trip.

 

Class meetings:      Lecture:           MWF 9-9:50 AM     (KGSX 217 or 309?)
                     Lab:                 Friday 1-4 PM         (KGSX 308)

                                   

Prerequisites:  BIO 369; CHM 105 or CHM 170 & 175

 

Required readings:

1.   Horne, A.J., and C.R. Goldman. 1994. Limnology. McGraw-Hill, 2nd edition.

2.   Reserve readings in library.

3.     Needham, J.G., and P.R. Needham. 1962. A guide to the study of freshwater biology.        Holden-Day, 5th edition.

4.     Havel, J.E. 2000. Limnology laboratory exercises. 7th edition. (Available at Textbucks,        located at Kimbrough & Madison).

Grading: Final grades  
Exam 1 50  90-100%  A
Exam 2 100  80-89 B
Exam 3 (final) 100 70-79  C
Laboratory participation 50 60-69 D
Laboratory reports 150 <60 F  
 450 points

 

Course policies

General philosophy.  Limnology is lab intensive and requires regular writing.  I will assume that students are familiar with the basic principles in the listed prerequisite courses.  Many students have difficulty with basic chemistry, so keep your general chemistry textbook handy.

Attendance.  Attendance in lecture will not be directly counted in grading for the course.  However, you are responsible for any announcements and handouts presented in your absence. Reading assignments are listed in the following schedule.  Attendance in laboratory is mandatory.  Each unexcused absence from lab will result in a loss of 20 points (participation) and three or more unexcused absences will result in an automatic failure for the course. Each absence from the Monday discussion period will result in the loss of five  points (participation).

Exams and makeup policy.  Three exams are scheduled.  Exams will be based on material from lecture, laboratory, and the text. I will provide study questions prior to each exam to guide your review of the text.  Examples of old exams are available on reserve; use these only as a guide for types of questions, not content.  If you must miss an exam, please let me know in advance. Makeup exams are usually all-essay format.

Laboratory, field trips, and discussion. The laboratory is vital part of this course.  The lab takes a lot of effort for us to set up, so there will be no opportunity to make up labs (but see “reports” below).  You are expected to help with cleanup after lab.  Since laboratory moves quickly, you should read the lab exercise prior to lab.  Scheduled field trips will go rain or shine, so you will need to dress appropriately for the weather. Although we will attempt to return to campus on time, Friday field trips may occasionally finish later than the scheduled period.  One field trip will take up an entire weekend, so adjust your schedule accordingly. By law, no alcohol is allowed in university vehicles. While off campus, remember that you are representing Missouri State, so common courtesy is expected.  The Monday lecture period will often be used to discuss the previous lab, so bring your lab data to this class.

Lab reports.  Lab reports are due on Monday, a week and a half after completion of the lab exercise.  Most reports are brief (10 points), though several other reports will be written in the format of scientific papers (20 points); details are given in the lab manual. For one excused absence, a 2-page summary on a pre-approved seminar can be substituted for a 10-point report and two such summaries can be substituted for a 20-point report. 

Academic integrity.  The following is general University policy: “All members of the University community share the responsibility and authority to challenge and make known acts of apparent academic dishonesty. Any student detected participating in any form of academic dishonesty in this course will be subject to sanctions as described in the Student Academic Integrity Policies and Procedures (http://www.missouristate.edu/acadaff/AcademicIntegrity.html).”  All lab reports or other written work must represent you own work.  We will share and interpret data during discussion, but all subsequent writing must be your own.  Any instances of copying will be considered academic dishonesty (score of zero for first infraction, failure of course for second infraction).

Disabilities.  Some of you may need accommodation for a physical or learning disability. Numerous campus resources are available through Disability Support Services (836-4192 or 6792 (TTY)).  If you have special needs, please visit with me during the first week of class.

Nondiscrimination.  Missouri State is an equal opportunity/affirmative action institution.  Concerns may be brought to the attention of your instructor (Havel), Biology Department Head (Jensen), or the University EO officer (Jana Long, 836-4252).

Library and internet resources

 

Reference books.  These books are all available at the Missouri State library.

APHA. 1989. Standard methods for the examination of water and wastewater. American Public Health Association. Washington, DC. 17th edition.

Downing, J.A., and F.H. Rigler. 1984.  A manual on methods for the assessment of secondary productivity in fresh waters. (IBP Handbook 17) Blackwell Scientific. Oxford, England.

Edmondson, W.T.(ed.) 1959.  Freshwater biology.  Wiley. New York. 2nd edition.

Hutchinson, G.E. 1957. A treatise on limnology. Volume 1: Lake origins, physics, and chemistry. Wiley.

Hutchinson, G.E. 1967. A treatise on limnology. Volume 2: Limnoplankton. Wiley.

Hutchinson, G.E. 1975. A treatise on limnology. Volume 3: Macrophytes. Wiley.

Hutchinson, G.E. 1993. A treatise on limnology. Volume 4: Benthos. Wiley.

Thorp, J.H., and A.P. Covich. 1991. Ecology and classification of North American freshwater invertebrates. Academic Press. New York

Welch, P.S.  1948.  Limnological methods.  McGraw Hill. New York.

Wetzel, R.G. 1983. Limnology. Saunders. New York. 2nd edition.  (On reserve)

Internet resources.  The aquatic biology web page contains a lot of helpful information for this class.  For example, we list library holdings for journals commonly used by limnologists and, in lecture, I will occasionally refer to web sites which are linked to the aquatic biology web page.  This web site can be found by first going to the Biology Department home page (http://biology.missouristate.edu/) and then clicking on “Aquatic Biology”.  A variety of useful web sites are available through the following link:  http://biology.missouristate.edu/aquatic/offcampuswebs.htm

Library reserve readings.  The following papers are required reading and are listed by title in the following lecture and lab schedules.  The papers are available through the library reserve, either by going to the desk and checking out a hard copy (3 hour limit) or by going to electronic reserve.  To use the electronic reserve, you may work from a computer lab on campus or else from your home computer.  From your home computer, you will need to first have Adobe Acrobat, which can be downloaded from the library web site.  Open the internet program, go to the Missouri State web site, locate the library web site, and then locate the reserves.  From reserves, search with my name (Havel) and then select BIO 562. You should then see a listing of reserve articles.  After choosing an article, give the following  password: limnology (all in lower case).  You should then see the text and can either read it on screen or else print and read later.  Note: for longer articles, printing is sometimes slow, so you might find visiting the reserve desk in person a little easier! 

Cole, G.A. 1994. “Shapes and sizes of lakes”. Pp. 132-150 in Textbook of limnology. Waveland Press, 4th edition.

Edmondson, W.T. 1991. “The eutrophication and recovery of Lake Washington”. Pp. 5-56 in The uses of ecology. University of Washington Press, Seattle.

Jones, J.R., and M.F. Knowlton. 1993. Limnology of Missouri reservoirs: An analysis of regional patterns. Lake and reservoir management 8: 17-30. (reference for lab report)

Korgen, B.J. 1995. Seiches. American Scientist 83: 330-341.

Lampert, W., and U. Sommer.  1997. “ Bacteria in freshwater”,  Pp. 93-100 in Limnoecology: The ecology of lakes and streams. Oxford University Press. New York.

Mallin, M.A.  2000. Impacts of industrial animal production on rivers and estuaries. American Scientist 88: 26-37.

Mason, C.F. 1991. “Sewage treatment”.  Pp. 57-72 in Biology of freshwater pollution. Longman Scientific. Singapore. 2nd edition.

Pechenik, J.A. 1987. “Writing laboratory reports”. Pp. 15-47 in Writing about biology. Little, Brown, and Co., Boston.

Wetzel, R.G. 1983. “Water economy”. Pp. 37-44 in Limnology. Saunders, 2nd edition.

 

 

Schedule
 

Jan        

19

Introduction (K. Pattinson)

ch.1

Limnology literature

Lab 1

   

   

   

   

   

   

Jan         

22

Lakes and ponds

ch. 2

  

   

    

24

Formation of lakes

pp. 457-464

   

   

26

Formation (continued)

          

Mapping and lake morphometry

Lab 2

   

  

   

  

  

   

Jan        

29

Sizes and shapes of lakes

Size and shape (Res.)

   

   

 

31

Light in water

ch. 3

   

   

Feb        

02

Heat and circulation

ch. 4

Heat models (long lab) **

Lab 3

   

  

   

   

   

   

Feb        

05

Discuss lab

---

   

   

  

07

Hydrology

Water economy (Res.)

    

   

   

09

Water movement

Seiches (Res.) 

Bioassays **

Lab 4

   

  

  

  

  

  

Feb       

12

EXAM 1 (50 points)

---

  

  

  

14

Bacteria (J. Steiert)

Bacteria in freshwater (Res.)

  

  

  

16

Oxygen (C. Barnhart)

ch. 7

NO LAB

---

  

  

  

  

  

  

Feb       

19

Presidents Day—HOLIDAY

---

  

  

  

21

Discuss exam and lab

---

  

  

  

23

Oxygen (cont.)

ch. 7

Sewage treatment plant 
(field trip)

Lab 5

  

  

  

  

  

  

Feb       

26

BOD and Discuss lab

Indus. Animal Prod. (Res.)

  

  

  

28

Carbon chemistry

ch. 7 & 6

  

  

Mar      

02

Acid rain

ch. 6

Water chemistry I

Lab 6

  

  

  

  

  

  

Mar      

05

Discuss lab

---

  

  

   

07

Nitrogen chemistry

ch. 8

  

  

  

09

Phosphorus chemistry

ch. 9

Water chemistry II **

Lab 7 & Limnology of Missouri reservoirs (Res.)

  

  

  

  

  

  

Mar      

12

Discuss lab

---

  

  

  

14

Chemistry CATCHUP

---

  

  

  

16

Biology of algae

pp. 200-204 & ch. 12

Algae diversity (R. Rhodes)

Lab 8, N&N pp.1-15, and 
web site

  

  

  

  

  

  

Mar      

19

Algae (continued)

pp. 200-204 
& ch. 12

  

  

  

21

Algae ecology

---

  

  

  

23

Biomass and productivity

pp. 350-354

Enumeration

Lab 9

  

  

  

  

  

  

Mar 

26-30

SPRING BREAK

---

NO LAB

---

  

  

  

  

  

  

Apr        

02

CATCHUP

---

  

  

04

EXAM 2 (100 points)

---

  

  

  

06

Eutrophication 

pp. 464-474 and Lake Wash. (Res.)

Zooplankton diversity

Lab 10, N&N pp. 17-27, and web site

  

  

  

  

  

  

Apr      

09

Eutrophication (cont.)

‘’         

  

  

  

11

Biology of zooplankton

Pp. 208-213, ch. 13

  

  

  

13

Spring holiday – NO CLASS

---

NO LAB

---

  

  

  

  

  

  

Apr       

16

Zooplankton: rotifers

Pp. 208-213, ch. 13

    

  

  

18

Zooplankton: cladocerans and copepods

          

  

  

  

20

Grazing, predation, and food webs

ch. 15

Limnological techniques (field trip)

Lab 11

  

  

  

  

  

  

Apr        

23

Discuss lab

---

  

  

   

25

Aquatic plants (macrophytes)

pp. 204-209

  

  

  

27

Littoral and benthic communities

pp. 213-216, 291-297

Bull Shoals Field Station
(weekend field trip)

Lab 12

  

  

  

  

  

  

Apr       

30

Stream ecology

ch. 16

  

  

May       

02

Stream ecology (cont.)

“       "

  

  

  

04

Rivers and their floodplains

ch. 17

BSFS group report

---

  

  

  

  

  

  

May      

07

Wetland ecology

ch. 18

NO LAB

---

  

09

CATCHUP

  

  

  

  

  

  

  

  

  

May     

17

FINAL EXAM (8:45 AM) (100 points)

  

  

  

* Readings for lecture:  text chapter or pages, article on reserve (Res.).  Readings for lab: chapter in lab manual (Lab), pages in field guide (N&N), or article on reserve.

**        Formal lab reports (20 points) for dates marked (**); abbreviated lab writeups (10 points) for other dates.

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Questions?   Comments? Contact Dr. Havel.

Copyright 1999 John E. Havel
This page last updated September 22, 2005.